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The Emperor Penguin (Aptenodytes forsteri) is the tallest and heaviest of all living penguin species and is endemic to Antarctica. The male and female are similar in plumage and size, reaching 122 cm (48 in) in height and weighing anywhere from 22 to 45 kg (49 to 99 lb). The dorsal side and head are black and sharply delineated from the white belly, pale-yellow breast and bright-yellow ear patches. Like all penguins it is flightless, with a streamlined body, and wings stiffened and flattened into flippers for a marine habitat. Its diet consists primarily of fish, but can also include crustaceans, such as krill, and cephalopods, such as squid. In hunting, the species can remain submerged up to 18 minutes, diving to a depth of 535 m (1,755 ft). It has several adaptations to facilitate this, including an unusually structured hemoglobin to allow it to function at low oxygen levels, solid bones to reduce barotrauma, and the ability to reduce its metabolism and shut down non-essential organ functions. The Emperor Penguin is perhaps best known for the sequence of journeys adults make each year in order to mate and to feed their offspring. The only penguin species that breeds during the Antarctic winter, it treks 50–120 km (31–75 mi) over the ice to breeding colonies which may include thousands of individuals. The female lays a single egg, which is incubated by the male while the female returns to the sea to feed; parents subsequently take turns foraging at sea and caring for their chick in the colony. The lifespan is typically 20 years in the wild, although observations suggest that some individuals may live to 50 years of age.

The Emperor Penguin's diet consists mainly of fish, crustaceans and cephalopods, although its composition varies from population to population. Fish are usually the most important food source, and the Antarctic silverfish (Pleuragramma antarcticum) makes up the bulk of the bird's diet. Other prey commonly recorded include other fish of the family Nototheniidae, the Glacial Squid (Psychroteuthis glacialis), and the hooked squid species Kondakovia longimana, as well as Antarctic krill (Euphausia superba).[45] The Emperor Penguin searches for prey in the open water of the Southern Ocean, in either ice-free areas of open water or tidal cracks in pack ice.One of its feeding strategies is to dive to around 50 m (164 ft), where it can easily spot sympagic fish like the Bald notothen (Pagothenia borchgrevinki) swimming against the bottom surface of the sea-ice; it swims up to the bottom of the ice and catches the fish. It then dives again and repeats the sequence about half a dozen times before surfacing to breathe.

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